篠山侏角龙属

篠山侏角龍屬; 化石时期：早白垩世，阿尔布阶 PreЄ Є O S D C P T J K Pg N
科学分类
界：	动物界 Animalia
门：	脊索动物门 Chordata
纲：	蜥形纲 Sauropsida
总目：	恐龍總目 Dinosauria
目：	†鸟臀目 Ornithischia
演化支：	†頭飾龍類 Marginocephalia
亚目：	†角龍亞目 Ceratopsia
下目：	†新角龍下目 Neoceratopsia
属：	†篠山侏角龍屬 Sasayamagnomus; Tanaka et al., 2024
模式種
	†三枝氏篠山侏角龙; Sasayamagnomus saegusai; Tanaka et al., 2024

篠山侏角龙（学名：Sasayamagnomus，意为“来自篠山的侏儒”）是新角龙类恐龙的一个属，来自日本兵库县早白垩世大山下组（英语：Ohyamashimo Formation）。属下包括单一物种三枝氏篠山侏角龙（S. saegusai）。^[1]估计正模标本长达80厘米（2.6英尺），尽管该标本尚未生长完全。^[2]

发现与命名

篠山侏角龙发现于兵库县丹波篠山市的宫田微脊椎动物地（Miyada microvertebrate site），已知化石包括17块脑颅骨骼、右喙骨及一个左胫骨。材料中保存的2个右鼻骨表明其中至少含2具个体。^[1]其正式描述之前曾在2023年的一篇会议摘要上公布。^[3]

其于2024年被正式描述为新角龙类新属新种。属名取自发现化石的篠山盆地及拉丁语单词“侏儒”。种名致敬丹波地区恐龙发掘活动的领导者，也是兵库县脊椎动物古生物学研究的卓越贡献者——三枝春雄（Haruo Saegusa）博士。^[1]

正模标本为MNHAH D1-060516。^[1]

分类

Tanaka等人将篠山侏角龙纳入系统发育分析，并发现其与鹰角龙及黎明角龙共处同一演化支。^[1]

古生态学

篠山侏角龙发现于大山下组（篠山群），后者年代为早白垩世末的阿尔布阶中早期。^[4]这些层位主要由砂岩、泥岩及砾岩构成。沉积环境（英语：depositional environment）代表具有半湿润（英语：Subhumid temperate climate）至半干旱气候的河流系统。^[5]恐龙牙齿在该层各地都很常见，包括属于其它兽脚类（驰龙科、镰刀龙下目及暴龙超科）及蜥脚类、甲龙类和禽龙类的牙齿。^[6]已描述了几个蛋化石類群（英语：ootaxa），包括姬蛋屬（英语：Himeoolithus）、纖長蛋屬（英语：Subtiliolithus）、日本蛋屬（英语：Nipponoolithus）及稜柱形蛋屬（英语：Prismatoolithus）。^[7]地层中还发现了怪蜥类（英语：Monstersauria）的双刃齿蜥、巨龙形类蜥脚下目丹波巨龙及伤齿龙科兽脚类寐猎龙。^[8]^[9]^[4]篠山群还有其它化石动物群，但原始描述中并未提及其发现于该地层，如真兽类的篠山兽（英语：Sasayamamylos）、^[10]石龙子亚目的厚颌蜥（英语：Pachygenys）^[11]及两种蛙类——兵库蛙（英语：Hyogobatrachus）与丹波蛙（英语：Tambabatrachus）。^[12]

参见

角龙类研究历史（英语：Timeline of ceratopsian research）

参考资料

^ ^1.0 ^1.1 ^1.2 ^1.3 ^1.4 Tanaka, Tomonori; Chiba, Kentaro; Ikeda, Tadahiro; Ryan, Michael J. A new neoceratopsian (Ornithischia, Ceratopsia) from the Lower Cretaceous Ohyamashimo Formation (Albian), southwestern Japan. Papers in Palaeontology. September 2024, 10 (5). doi:10.1002/spp2.1587.
^ 兵庫県で発見の角竜類の化石新属新種と確認 | お知らせ・トピックス. 岡山理科大学. [2024-09-04] （日语）.
^ Tanaka, Tomonori; Chiba, Kentaro; Ikeda, Tadahiro; Kubota, Katsuhiro. Phylogenetic position of a ceratopsian dinosaur from the Lower Cretaceous Ohyamashimo Formation (Albian) of the Sasayama Group in Tambasasayama City, Hyogo, Japan (PDF). The Palaeontological Society of Japan. 2023.
^ ^4.0 ^4.1 Kubota, K.; Kobayashi, Y.; Ikeda, T. Early Cretaceous troodontine troodontid (Dinosauria: Theropoda) from the Ohyamashimo Formation of Japan reveals the early evolution of Troodontinae. Scientific Reports. 2024, 14. 16392. PMC 11272788 . doi:10.1038/s41598-024-66815-2 .
^ Hayashi, Keiichi; Fujita, Saki; Koarai, Kazuto; Matsukawa, Masaki. Stratigraphy and paleoenvironment of the Cretaceous Sasayama Group in the Sasayama area, Hyogo Prefecture, western Japan. The Journal of the Geological Society of Japan. 2017-09-15, 123 (9): 747–764. ISSN 0016-7630. doi:10.5575/geosoc.2017.0016  （英语）.
^ Kubota, Katsuhiro. 日本産の中生代恐竜化石目録 2022年版 [A list of Mesozoic dinosaur fossils from Japan in 2022] (PDF). Bulletin of the Gunma Museum of Natural History: 157–170.
^ Tanaka, Kohei; Zelenitsky, Darla K.; Therrien, François; Ikeda, Tadahiro; Kubota, Katsuhiro; Saegusa, Haruo; Tanaka, Tomonori; Ikuno, Kenji. Exceptionally small theropod eggs from the Lower Cretaceous Ohyamashimo Formation of Tamba, Hyogo Prefecture, Japan. Cretaceous Research. 2020, 114: Article 104519. Bibcode:2020CrRes.11404519T. S2CID 219449961. doi:10.1016/j.cretres.2020.104519.
^ Ikeda, T.; Ota, H.; Tanaka, T.; Ikuno, K.; Kubota, K.; Tanaka, K.; Saegusa, H. A fossil Monstersauria (Squamata: Anguimorpha) from the Lower Cretaceous Ohyamashimo Formation of the Sasayama Group in Tamba City, Hyogo Prefecture, Japan. Cretaceous Research. 2021, 130: Article 105063. S2CID 239230916. doi:10.1016/j.cretres.2021.105063 .
^ Saegusa, H.; Ikeda, T. A new titanosauriform sauropod (Dinosauria: Saurischia) from the Lower Cretaceous of Hyogo, Japan. Zootaxa. 2014, 3848 (1): 1–66. PMID 25112425. doi:10.11646/zootaxa.3848.1.1 .
^ Kusuhashi, Nao; Tsutsumi, Yukiyasu; Saegusa, Haruo; Horie, Kenji; Ikeda, Tadahiro; Yokoyama, Kazumi; Shiraishi, Kazuyuki. A new Early Cretaceous eutherian mammal from the Sasayama Group, Hyogo, Japan. Proceedings of the Royal Society B: Biological Sciences. 2013-05-22, 280 (1759): 20130142. ISSN 0962-8452. PMC 3619506 . PMID 23536594. doi:10.1098/rspb.2013.0142 （英语）.
^ Ikeda, Tadahiro; Ota, Hidetoshi; Saegusa, Haruo. A new fossil lizard from the Lower Cretaceous Sasayama Group of Hyogo Prefecture, western Honshu, Japan. Journal of Vertebrate Paleontology. 2015-01-02, 35 (1): e885032. ISSN 0272-4634. doi:10.1080/02724634.2014.885032 （英语）.
^ Ikeda, Tadahiro; Ota, Hidetoshi; Matsui, Masafumi. New fossil anurans from the Lower Cretaceous Sasayama Group of Hyogo Prefecture, Western Honshu, Japan. Cretaceous Research. 2016-06-01, 61: 108–123. ISSN 0195-6671. doi:10.1016/j.cretres.2015.12.024.

[Sasayamagnomus-1] 1.0 ^1.1 ^1.2 ^1.3 ^1.4 Tanaka, Tomonori; Chiba, Kentaro; Ikeda, Tadahiro; Ryan, Michael J. A new neoceratopsian (Ornithischia, Ceratopsia) from the Lower Cretaceous Ohyamashimo Formation (Albian), southwestern Japan. Papers in Palaeontology. September 2024, 10 (5). doi:10.1002/spp2.1587.

[2] 兵庫県で発見の角竜類の化石新属新種と確認 | お知らせ・トピックス. 岡山理科大学. [2024-09-04] （日语）.

[PSJ2023_ceratops-3] Tanaka, Tomonori; Chiba, Kentaro; Ikeda, Tadahiro; Kubota, Katsuhiro. Phylogenetic position of a ceratopsian dinosaur from the Lower Cretaceous Ohyamashimo Formation (Albian) of the Sasayama Group in Tambasasayama City, Hyogo, Japan (PDF). The Palaeontological Society of Japan. 2023.

[Hypnovenator-4] 4.0 ^4.1 Kubota, K.; Kobayashi, Y.; Ikeda, T. Early Cretaceous troodontine troodontid (Dinosauria: Theropoda) from the Ohyamashimo Formation of Japan reveals the early evolution of Troodontinae. Scientific Reports. 2024, 14. 16392. PMC 11272788 . doi:10.1038/s41598-024-66815-2 .

[SasayamaStrat-5] Hayashi, Keiichi; Fujita, Saki; Koarai, Kazuto; Matsukawa, Masaki. Stratigraphy and paleoenvironment of the Cretaceous Sasayama Group in the Sasayama area, Hyogo Prefecture, western Japan. The Journal of the Geological Society of Japan. 2017-09-15, 123 (9): 747–764. ISSN 0016-7630. doi:10.5575/geosoc.2017.0016  （英语）.

[List2022-6] Kubota, Katsuhiro. 日本産の中生代恐竜化石目録 2022年版 [A list of Mesozoic dinosaur fossils from Japan in 2022] (PDF). Bulletin of the Gunma Museum of Natural History: 157–170.

[Tanaka2020-7] Tanaka, Kohei; Zelenitsky, Darla K.; Therrien, François; Ikeda, Tadahiro; Kubota, Katsuhiro; Saegusa, Haruo; Tanaka, Tomonori; Ikuno, Kenji. Exceptionally small theropod eggs from the Lower Cretaceous Ohyamashimo Formation of Tamba, Hyogo Prefecture, Japan. Cretaceous Research. 2020, 114: Article 104519. Bibcode:2020CrRes.11404519T. S2CID 219449961. doi:10.1016/j.cretres.2020.104519.

[Morohasaurus-8] Ikeda, T.; Ota, H.; Tanaka, T.; Ikuno, K.; Kubota, K.; Tanaka, K.; Saegusa, H. A fossil Monstersauria (Squamata: Anguimorpha) from the Lower Cretaceous Ohyamashimo Formation of the Sasayama Group in Tamba City, Hyogo Prefecture, Japan. Cretaceous Research. 2021, 130: Article 105063. S2CID 239230916. doi:10.1016/j.cretres.2021.105063 .

[Tambatitanis-9] Saegusa, H.; Ikeda, T. A new titanosauriform sauropod (Dinosauria: Saurischia) from the Lower Cretaceous of Hyogo, Japan. Zootaxa. 2014, 3848 (1): 1–66. PMID 25112425. doi:10.11646/zootaxa.3848.1.1 .

[10] Kusuhashi, Nao; Tsutsumi, Yukiyasu; Saegusa, Haruo; Horie, Kenji; Ikeda, Tadahiro; Yokoyama, Kazumi; Shiraishi, Kazuyuki. A new Early Cretaceous eutherian mammal from the Sasayama Group, Hyogo, Japan. Proceedings of the Royal Society B: Biological Sciences. 2013-05-22, 280 (1759): 20130142. ISSN 0962-8452. PMC 3619506 . PMID 23536594. doi:10.1098/rspb.2013.0142 （英语）.

[11] Ikeda, Tadahiro; Ota, Hidetoshi; Saegusa, Haruo. A new fossil lizard from the Lower Cretaceous Sasayama Group of Hyogo Prefecture, western Honshu, Japan. Journal of Vertebrate Paleontology. 2015-01-02, 35 (1): e885032. ISSN 0272-4634. doi:10.1080/02724634.2014.885032 （英语）.

[12] Ikeda, Tadahiro; Ota, Hidetoshi; Matsui, Masafumi. New fossil anurans from the Lower Cretaceous Sasayama Group of Hyogo Prefecture, Western Honshu, Japan. Cretaceous Research. 2016-06-01, 61: 108–123. ISSN 0195-6671. doi:10.1016/j.cretres.2015.12.024.

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